Peer-reviewed Articles

Gómez, R.A., Will, K.W. and Maddison, D.R. (2016). Are Miquihuana rhadiniformis Barr, 1982 and Pseudamara arenaria (LeConte, 1847) (Coleoptera, Carabidae) sphodrines? Phylogenetic analysis of data from next-generation sequencing of museum specimens resolves the tribal-group relationships of these enigmatic taxa. Entomologische Blätter und Coleoptera (2016) 112(1):149–168.

Will, K.W. and Steplowski, I. 2016. A 3D printed Malaise trap head. Pan-Pacific Entomologist. 92(2):86–91.

Gómez, R.A., Reddell, J., Will, K.W. and Moore, W. 2016. Up high and down low: Molecular systematics and insight into the diversification of the ground beetle genus Rhadine LeConte. Molecular Phylogenetics and Evolution 98, 161–175.

Will, K.W. 2015. A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa, 4034(2), pp.291–308.

Will, K.W. 2015. Resolution of taxonomic problems in Australian Harpalini, Abacetini, Pterostichini, and Oodini (Coleoptera, Carabidae). ZooKeys, 545, pp.131–137.

Will, K.W. 2015. A multigene phylogenetic analysis results in a redefinition of the genus Notonomus Chaudoir (Coleoptera, Carabidae) and descriptions of new species of the subgenus Castelnau. Invertebrate Systematics, 29:4.

Liebherr, J.K. and Will, K.W. 2015. Antisymmetric male genitalia in Western Australian populations of Mecyclothorax punctipennis (Coleoptera: Carabidae: Moriomorphini). Insect Systematics & Evolution, 1–17.

Grzymala, T. and Will, K.W. 2014. Taxonomic review of Cratocerus Dejean, 1829 (Coleoptera, Carabidae) with the description of six new species. ZooKeys 416, 77–112.

Will, K.W. and Kavanaugh, D.H. 2012. A new species of Lesticus Dejean, 1828 (Coleoptera, Carabidae) from the Finisterre Range, Papua New Guinea and a key to the genera of pterostichine-like Harpalinae of New Guinea. ZooKeys, (246), 27–37.

Will, K.W. 2011. Taxonomic review of the Pterostichini and Loxandrini fauna of New Caledonia (Coleoptera, Carabidae). ZooKeys, (147), 337–97.

Will, K.W. 2011. Chaetauchenium Tschitchérine, 1900 ( Carabidae: Pterostichini ) a new species and new status for a lineage from the Valdivian temperate rainforest. Gayana, 75(2), 192–197.

Guthrie, N.A., Weir, T. and Will, K.W. 2010. Localised and regional patterns in ground-dwelling beetle assemblages in a semi-tropical arid zone environment. Records of the Western Australian Museum Supplement 78, pp. 169–184.

Will, K.W. 2010. Synonymy in Sugimotoa Habu (Coleoptera: Carabidae: Lebiini). The Coleopterists Bulletin, 64(1), 91.

Will, K.W., Gill, A.S., Lee, H. and Attygalle, A.B. 2010. Quantification and evidence for mechanically metered release of pygidial secretions in formic acid-producing carabid beetles. Journal of Insect Science (Online), 10(12)17pp.

Ruiz, C., Jordal, B.H., Emerson, B.C., Will, K.W. and Serrano, J. 2010. Molecular phylogeny and Holarctic diversification of the subtribe Calathina (Coleoptera: Carabidae: Sphodrini). Molecular Phylogenetics and Evolution, 55(2), 358–71.

Attygalle, A., Wu, X., Maddison, D.R. and Will, K.W. 2009. Limonene and other monoterpenes in the defensive secretion of some carabid beetles. Natuwissenschaften 96(12):1443.

Anichtchenko, A. and Will, K.W. 2009. A new species of Stolonis Motschulsky, 1866 (Coleoptera, Carabidae) from Argentina. Russian Entomological Journal 18(1):51–53.

Will, K.W. 2008. A new species of Loxandrus LeConte (Coleoptera: Carabidae: Loxandrini) from South America. Annals of Carnegie Museum 77(1)205–210.

Will, K.W. and Gill, A.S. 2008. Phylogeny and classification of Hypherpes auctorum (Coleoptera: Carabidae: Pterostichini: Pterostichus). Annals of Carnegie Museum 77(1)93–127.

Will, K.W. and Park, J.K. 2008. A new Species of the oriental Abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus. The Coleopterists Bulletin 62(2)189–196.

Park, J.K. and Will, K.W. 2008. The first records of Zuphiini (Coleoptera: Carabidae) from New Caledonia; A new genus and two new species. Memoirs of the Queensland Museum 52(2): 99–104.

Will, K.W. 2007. Four New Species of the subgenus Leptoferonia Casey (Coleoptera, Carabidae, Pterostichus Bonelli) from California. Proceedings of the California Academy of Sciences 58(4)49–57.

Attygalle, A.B., Wu, X. and Will, K.W. 2007. Biosynthesis of tiglic, ethacrylic, and 2-methylbutyric acids in a carabid beetle, Pterostichus (Hypherpes) californicus. Journal of Chemical Ecology 33:963–970.

Rubinoff, D., Cameron, S. and Will, K.W. 2006. A Genomic perspective on the shortcomings of mitochondrial DNA for Barcoding. Journal of Heredity 97(6)581–594.

Park, J.K., Dam, H.T. and Will, K.W. 2006. Carabidae from Vietnam (Coleoptera). Journal of Asia-Pacific Entomology 9(2):1–61.

Cameron, S. Rubinoff, D. and Will, K.W. 2006. Who will actually use DNA barcoding and what will it cost? Systematic Biology 55(5):844–847.

Will, K.W. 2006. A new species of the New Caledonian genus Sphodrosomus Perroud (Coleoptera: Carabidae: Pterostichini). Memoirs of the Queensland Museum 52(1) 281–285.

Rubinoff, D., Cameron, S. and Will, K.W. 2006. Are plant DNA barcodes a search for the Holy Grail? Trends in Ecology and Evolution 21(1):1–2.

Will, K.W. and Liebherr, J.K. 2005. Removal of Hannaphota distincta Landin (Coleoptera: Carabidae) from Pterostichini to Platynini. The Coleopterists Bulletin 59(4): 448–449.

Will, K.W, Mishler, B.D. and Wheeler, Q.D. 2005. The perils of DNA barcoding and the need for integrative taxonomy. Systematic Biology 54(5):844–851.

Will, K.W. 2005. New tribal and generic placement for taxa of Pterostichini (auct.) (Coleoptera: Carabidae) from the Juan Fernández Archipelago, Chile with taxonomic notes on Trirammatus Chaudoir, 1838. The Pan-Pacific Entomologist 81(1/2):68–75.

Will, K.W. 2005. The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini). Zootaxa 1049:1–17.

Will, K.W., Liebherr, J.K., Maddison D.R. and Galián, J. 2005. Absence asymmetry: the evolution of monorchid beetles (Insecta:Coleoptera: Carabidae). Journal of Morphology 264:75–93.

Will, K.W. 2004. A remarkable new species of Trirammatus Chaudoir (Coleoptera: Carabidae: Pterostichini) from the Valdivian forest of Chile. Zootaxa 758: 1–9.

Will, K.W. 2005. New species of the South American loxandrine genus Metoncidus Bates (Coleoptera: Carabidae). Revista Brasileira de Entomologia 48(4):441–446.

Will, K.W. 2005. A new genus of Caelostomini (Coleoptera: Carabidae) from Papua New Guinea. Coleopterists Bulletin 58(4):637–641.

Will, K.W. 2005. Review and cladistic analysis of the generic-level taxa of Morionini Brullé‚ (Coleoptera: Carabidae). The Pan-Pacific Entomologist 79(3/4):212–229.

Will, K.W. and Rubinoff, D. 2004. Myth of the molecule: DNA barcodes for species cannot replace morphology for identification and classification. Cladistics 20: 47–55.

Attygalle, A.B., Wu, X., Ruzicka, J., Rao, S., Garcia, S., Herath, K., Meinwald, J., Maddison, D.R. and Will, K.W. 2004. Defensive chemicals of two species of Trachypachus Motschulski. Journal of Chemical Ecology 30: 577–588.

Will, K.W. 2002. Revision of the new world abariform genera Neotalus n.gen. and Abaris Dejean (Coleoptera: Carabidae: Pterostichini (auctorum). Annals of the Carnegie Museum of Natural History 71:143–213.

Will, K.W. and Liebherr, J.K. 2002. A new tribal placement for Glyptolenus mirabilis (Straneo) (Coleoptera: Carabidae), comb. n., and description of G. straneoi, sp. n. from Ecuador. Studies on Neotropical Fauna and Environment 37:59–64.

Will, K.W., Attygalle, A.B. and Herath, K. 2001. Erratum, new defensive chemical data for ground beetles (Coleoptera: Carabidae): interpretations in a phylogenetic framework. Biological Journal of the Linnean Society 73:167.

Will, K.W. 2001. A new species of Harpalus Latreille (Coleoptera: Carabidae) from southeastern North America. The Coleopterists Bulletin 55(4):447–452.

Will, K.W., Attygalle, A.B. and Herath, K. 2000. New defensive chemical data for ground beetles (Coleoptera: Carabidae): interpretations in a phylogenetic framework. Biological Journal of the Linnean Society 71:459–481.

Marshall, C.J. and Will, K.W. 2000. Should type specimens of insects, indigenous to one country but housed in another, be returned to their country of origin? -Background Statement. In: ed. L.L. Allee, International issues in entomology: a student debate. American Entomologist 46(3):166–167.

Will, K.W. 1998. Oviposition behavior in Dromius piceus Dejean (Coleoptera: Carabidae, Lebiini). Fabreries 23(3-4):120–123.

Will, K.W. 1998. A new species of Diplocheila Brulle‚ from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini). Proceedings of the Entomological Society of Washington 100(1):95–103.

Samuelson, G.A., Liebherr, J.K. and Will, K.W. 1997. Gnathaphanus picipes, an established adventive in Hawaii (Coleoptera: Carabidae). Bishop Museum Occasional Papers 49(2):22–23.

Will, K.W. and Liebherr, J.K. 1997. New and little known species of Loxandrus LeConte 1852 (Coleoptera: Carabidae) from North and South America. Studies on Neotropical Fauna and Environment 32(4):230–238.

Will, K.W. 1997. Review of the species of the subgenus Megapangus Casey (Coleoptera: Carabidae: Harpalini: Harpalus (Latreille). The Coleopterists Bulletin 51(1):43–51.

Liebherr, J.K. and Will, K.W. 1996. A new Pseudomorpha Kirby (Coleoptera: Carabidae: Pseudomorphini) from the Sierra de los Tuxtlas, Veracruz, Mexico. Folia Entomologica Mexicana 98:53–58.

Liebherr, J.K. and Will, K.W. 1996. New North American Platynus Bonelli (Coleoptera: Carabidae), a key to species north of Mexico and notes on species from the southwestern United States. The Coleopterists Bulletin 50(4):301–320.

Will, K.W., Purrington, F.F. and Horn, D.J. 1995. Ground beetles of islands in the western basin of Lake Erie and the adjacent mainland (Coleoptera: Carabidae, including Cicindelini). The Great Lakes Entomologist 28(1):55–70.

Will, K.W. 1995. Overwintering of Pollenia rudis (Diptera: Calliphoridae). Entomological News 106(4):177.

Will, K.W. 1995. Plecoptera surface-skimming and insect flight evolution. Science 270(5242):1684–1685.

Contributed papers to edited and reviewed volumes

Elgueta, M., Arias, E. and Will, K.W. 2008. Curculionoidea (Coleoptera) en follaje de arboles del centro-sur de Chile. In: J.E. Llorente and A. Lanteri (Eds), Contribuciones Taxonómicas en Órdenes de Insectos Hiperdiversos. Universidad Nacional Autónoma de México, México, D.F., pp. 177–200.

Liebherr, J.K. and Will, K.W. 1998. Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract. Pp 107–170. in Phylogeny and Classification of Caraboidea Coleoptera: Adephaga). In: G.E. Ball, A. Casale, and A.V. Taglianti (Eds), Proceedings of a Symposium (28 August, 1996. Florence, Italy). XX International Congress of Entomology. Atti Museo Regionale di Scienze Naturali di Torino. Italy, pp. 107–170.

Will, K.W. 1999. Systematics and zoogeography of the genus Lophoglossus LeConte (Coleoptera Carabidae Pterostichini). In: A. Zamotajlov and R. Sciaky (Eds), Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij. MUISO Publishers, Krasnodar, pp. 259–276.

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