Tortricidae

Classification following Horak (1999) Handbook of Zoology

.Tortricinae
......Phricanthini
......Tortricini
......Schoenotenini
......Cochylini
......Cnephasiini
......Archipini
......Sparganothini
......Atteriini
......Euliini
......Ceracini
.Chlidanotinae
......Polyorthini
......Chlidanotini
......Hilarographini
.Olethreutinae
......Microcorsini
......Gatesclarkeanini
......Bactrini
......Olethreutinae
......Enarmoniini
......Eucosmini
......Grapholitini

Tahitian Dichelopa
Dichelopa sp. from Tahiti (Mt. Aorai), Society Islands, French Polynesia, PT Oboyski 2002

 

Grapholitini

Phylogeny of Palaearctic (& Australian) Grapholitini by Komai 1999 (modified Komai & Horak 2006)
Komai Phylogeny
For exceptions to characters and states see:
Komai F. 1999. A taxonomic review of the genus Grapholita and allied genera (Lepidoptera: Tortricidae) in the Palaearctic region. Entomologica Scandinavica, Supplement 55: 4-226

 

Phylogeny of Palaearctic Grapholitini by Danilevsky & Kuznetsov (1968)
Danilevsky Kuznetsov phylogeny
The original phylogeny is a hand-drawn tree (literally) that gives more detail regarding time of splitting and branch "thickness" (see pg. 69 in D & K 1968, reproduced on pg. 35 in Komai 1999).
For details see:
Danilevsky AS & Kuznetsov VI. 1968. Tortrix Moths Tortricidae: Tribe Laspeyresiini. Fauna USSR, Lepidoptera, Serial 7, 5(1):1-635 (in Russian)

 

 Phylogeny of North American Grapholitini by Heinrich (1926)
Heinrich Phylogeny
Notes: Laspeyresia, Melissopus, and Carpocapsa have been synonymized as Cydia since Heinrich's publication. In Heinrich's original phylogeny, Ethelgoda, Sereda, and Grapholita directly derive from Laspeyresia (rather than Laspeyresia being sister). Likewise, Satronia derives from Ricula. Heinrich's phylogeny is not a parsimony-based phylogeny based on the characters listed in the "Comparative table of structural characters: Laspeyresiinae)." Only four characters (1-4 above) have any phylogenetic signal. A heuristic parsimony analysis based on these characters (and synonymizing the Cydia genera) results in two most parsimonious trees (roll over text above to view, or click to open in a new window).
For exceptions and further explanations of characters and states see:
Heinrich C. 1926. Revision of the North American moths of the subfamilies Laspeyresiinae and Olethreutinae. US National Museum, Bulletin 132: 1-75