Leathers, K., Herbst, D., Safeeq, M. & A. Ruhi. Dynamic, downstream-propagating thermal vulnerability in a mountain stream network: implications for biodiversity in the face of climate change. Limnology and Oceanography, in press.

Fournier, R., de Mendoza, G., Sarremejane, R. & A. Ruhi (2022). Isolation controls reestablishment mechanisms and post-drying community structure in an intermittent stream. Ecology, accepted for publication.

Ruhi, A., Hwang, J., Devineni, N., Mukhopadhyay, S., Kumar, H., Comte, L., Worland, S. &  A. Sankarasubramanian (2022). How does flow alteration propagate across a large, highly-regulated basin? Dam attributes, network context, and implications for biodiversity. Earth’s Future, 10(6): e2021EF002490.

Colombano, D., Carlson, S.M., Hobbs, J. & A. Ruhi (2022). Four decades of climatic fluctuations and fish recruitment stability across a marine-freshwater gradient. Global Change Biology, 28(17): 5104-5120.

Baruch, E.M., Ruhi, A., Harms, T.K. & J.L. Sabo (2022). Flow variation at multiple scales filters fish life histories and constrains community diversity in desert streams. Ecosphere, 13(6): e4086.

Mor, J.R, Muñoz, I., Sabater, S., Zamora, L. & A. Ruhi (2022). Energy limitation or sensitive predators? Trophic and non-trophic impacts of wastewater pollution on stream food webs. Ecology, 103(2), e03587.


Hwang, J., Kumar, H., Ruhi, A., Sankarasubramanian, A. and Devineni, N., 2021. Quantifying Dam‐Induced Fluctuations in Streamflow Frequencies Across the Colorado River Basin. Water Resources Research, 57(10), p.e2021WR029753.

Comte, L., Olden, J.D., Tedesco, P., Ruhi, A & X. Giam (2021). Climate and land-use changes interact to drive long-term reorganization of riverine fish communities globally. Proceedings of the National Academy of Sciences of the United States of America, 118(27).

Sarremejane, R., Stubbington, R., England, E., Sefton, C.E.M., Eastman, M., Parry, S. & A. Ruhi (2021). Drought effects on invertebrate metapopulation dynamics and quasi-extinction risk in an intermittent river network. Global Change Biology, 27(17): 4024–4039.

Chalise, D.R., Sankarasubramanian, A. & A. Ruhi (2021). Dams and climate interact to alter river flow regimes across the United States. Earth’s Future, 9(4), p.e2020EF001816.

Benito. X ., Luethje, M., Schneider, T., Fritz, S.C., Baker, P., Pedersen, E., Gauzere, P., Nascimiento, M., Bush, M. & A. Ruhi (2021). Ecological resilience in tropical Andean lakes: a paleolimnological perspective. Limnology & Oceanography, 67: S23–S37.

Larsen, L., Comte L., Filipe A.F., Fortin M.-J., Jacquet C., Remo Ryser, Tedesco P.A., Brose U., Erős T., Giam X., Irving K., Ruhi A., Sharma S., Olden J.D (2021). The geography of metapopulation synchrony in dendritic river networks. Ecology Letters, 24: 791–801.

Patrick, C.J., McCluney, K.E., Ruhi, A., Gregory, A., Sabo, J. & J.H. Thorp (2021). Multi-scale biodiversity drives temporal variability in macrosystems. Frontiers in Ecology and the Environment, 19(1):47-56.

Vander Vorste, R, Stubbington, R., Acuña, V., Bogan, M.T., Bonada, N., Cid. N., Datry, T., Storey, R., Wood, P.J. & A. Ruhi (2021). Climatic aridity increases temporal nestedness of invertebrate communities in naturally drying rivers. Ecography, 44: 1-10.

Comte, L., Grantham, T. & A. Ruhi (2021). Human stabilization of river flows is linked with fish invasions across the United States. Global Ecology and Biogeography, 30(3):725–737.

Comte, L., et al.* (2021). RivFishTIME: A global database of fish time-series to study global change ecology in riverine systems. Global Ecology and Biogeography, 30(1):38-50. [*more than 30 co‑authors].


Sanchez-Montoya, M., Tockner, K., von Schiller, D., Miñano, J., Catarineu, C., Lencina, J.L., Gonzalez-Barbera, G. &. A. Ruhi (2020). Dynamics of ground-dwelling arthropod metacommunities in intermittent streams: the key role of dry riverbeds. Biological Conservation 241:108328.

Allen, D., Datry, T., Boersma, K., Bogan, M., Boulton, A.J., Bruno, D., Busch, M., Costigan, K., Dodds, W.K., Fritz, K., Godsey, S.E., Jones, J.B., Kaletova, T., Kampf, S.K., Mims, M.C., Neeson, T.M., Olden, J.D., Pastor, A.V., Poff, N.L. Ruddell, B.L., Ruhi, A., Singer, G., Vezza, P., Ward, A.S. & M. Zimmer (2020).  River ecosystem conceptual models and non-perennial rivers: A critical review. Wiley Interdisciplinary Reviews: Water, 7(5):e1473.

Erős, T., Comte, L., Filipe, A.F., Ruhi, A., Tedesco, P.A., Brose, U. Fortin, M.J., Giam, X., Irving, K., Jacquet, C., Larsen, S., Sharma, S. & J.D. Olden (2020). Effects of nonnative species on the stability of riverine fish communities. Ecography, 43(8): 1156-1166.


Palmer, M & A. Ruhi (2019). Linkages between flow regime, biota, and ecosystem processes: implications for river restoration. Science 365, eaaw2087.

Anderson, E.P., Jackson, S., Tharme, R.E., Douglas, M., Flotemersch, J.E., Zwarteveen, M., Lokgariwar, C., Montoya, M., Wali, A., Tipa, G.T. Jardine, T.D., Olden, J.D., Cheng, L., Conallin, J., Cosens, B., Dickens, C., Garrick, D., Groenfeldt, D., Kabogo, J., Roux, D., Ruhi, A. & A.H. Arthington (2019). Understanding rivers and their social relations: A critical step to advance environmental water management. Wiley Interdisciplinary Reviews: Water, p.e1381.

Tonkin, J.D., Poff, N.L., Bond, N.R., Horne, A., Merritt, D.M., Reynolds, L.V., Olden, J.D., Ruhi, A. & Lytle, D.A., 2019. Prepare river ecosystems for an uncertain future. Nature 570 (7761):301.

Sabo, J.L., Holtgrieve, G.W., Ruhi, A., Arias, M.E., Ngor, P.B., Elliott, V., Räsänen, T.A. & S. Nam (2019). Response to Comment on “Designing river flows to improve food security futures in the Lower Mekong Basin”. Science 364(6444): eaav9887.

Ruhi, A., Cross, W., Catford, J., Escoriza, D. & J.D. Olden (2019). Understanding the nexus between hydrological alteration and biological invasions. In: Sabater, S., Elosegi, A. & R. Ludwig (Eds), Multiple Stressors in River Ecosystems. Elsevier.


Palmer, M. & A. Ruhi (2018). Measuring Earth’s rivers. Science 361(6402): 546-547.

Ruhi, A., Messager, M. & J.D. Olden (2018). Tracking the pulse of the Earth’s fresh waters. Nature Sustainability 1:198–203.

Ruhi, A., Dong, X., Holt, C., Batzer, D.P. & J.L. Sabo (2018). Detrimental effects of a novel flow regime on the functional trajectory of an aquatic invertebrate metacommunity. Global Change Biology 24(8): 3749-3765.

Holtgrieve, G.W., Arias, M.E., Ruhi, A., Elliott, V., Nam, S., Ngor, P.B., Räsänen, T.A.  & J.L. Sabo (2018). Response to Comments on “Designing river flows to improve food security futures in the Lower Mekong Basin”. Science 361(6398): p.eaat1477.

Sabo, J.L., Caron, M., Doucett, R., Dibble, K., Ruhi, A., Marks, J.C., Hungate, B & T.A. Kennedy (2018). Pulsed flows, tributary inputs, and food-web structure in a highly regulated river. Journal of Applied Ecology 55(4):1884-1895.

Kominoski, J.S., Ruhi, A., Hagler, M.M., Petersen, K., Sabo, J.L., Sinha, T., Arumugam, S. & J.D. Olden (2018). Patterns and drivers of fish extirpations in rivers of the American Southwest and Southeast. Global Change Biology 24(3):1175–1185.

Mor, J.R., Ruhi, A., Muñoz, I., Tornés, E., Acuña, V., & S. Sabater (2018). Dam regulation and riverine food-web structure in a Mediterranean river. Science of the Total Environment 625:301-310.

2017 and older

Ruhi, A., Datry, T. & J.L. Sabo (2017). Interpreting beta diversity components over time to conserve metacommunities in highly-dynamic ecosystems. Conservation Biology 31(6):1459-1468.

Sabo, J.L., Ruhi, A., Elliot, V., Arias, M., Holtgrieve, G. & N. So (2017). Designing river flows to improve food security futures in the Lower Mekong Basin. Science 358(6368): eaao1053. [cover article]

Sankarasubramanian, A., Sabo, J. L., Larson, K., Seo, S., Sinha, T., Bhowmik, R., Ruhi, A., Kunkel, K., Mahinthakumar, G., Berglund, E. & J. Kominoski (2017). Synthesis of public water supply use in the U.S.: Spatio-temporal patterns and socio-economic controls. Earth’s Future 5(7):771-788.

Dong, X., Ruhi, A. & N.B. Grimm (2017). Evidence for self-organization in determining spatial patterns of stream nutrients, despite primacy of the geomorphic template. Proceedings of the National Academy of Sciences of the United States of America 114(24):4744–4752.

Rosset, B., Ruhi, A., Bogan, M.T. & T. Datry (2017). Do lentic and lotic communities respond similarly to drying? Ecosphere 8(7):e01809.

Acuña, V., Hunter, M. & A. Ruhi (2017). Managing temporary streams and rivers as unique rather than second-class ecosystems. Biological Conservation 211(B):12-19.

Bogan, M.T., Chester, E., Datry, T., Murphy, A., Robson, B., Ruhi, A. & J. Whitney (2017). Resistance, Resilience, and Community Recovery in Intermittent Rivers and Ephemeral Streams. In: Datry, T., Bonada, N. & A. Boulton (Eds), Intermittent Rivers and Ephemeral Streams: Ecology and Management. Elsevier.

Datry, T., Bonada, N., Corti, R., Heino, J., Hugueny, B., Rolls. R & A. Ruhi (2017). Habitat Fragmentation and Metapopulation, Metacommunity, and Metaecosystem Dynamics in Intermittent Rivers and Ephemeral Streams. In: Datry, T., Bonada, N. & A. Boulton (Eds.), Intermittent Rivers and Ephemeral Streams: Ecology and Management. Elsevier.

Ruhi, A., Olden, J. & J.L. Sabo (2016). Declining streamflow induces collapse and replacement of native fish in the American Southwest. Frontiers in Ecology and the Environment 14(9):1-8. [F1000 Prime recommended]

Ruhi, A., Muñoz, I., Tornés, E., Batalla, R.J., Vericat, D., Ponsatí, L., Acuña, V., von Schiller, D., Marcé, R., Bussi, G., Francés, F. & S. Sabater (2016). Flow regulation increases foodchain length through omnivory mechanisms in a Mediterranean river network. Freshwater Biology 61(9):1536-1549.

Ruhi, A., Acuña, V., Barceló, D., Huerta, B., Mor, J.R., Rodríguez, S. & S. Sabater (2016). Bioaccumulation and trophic magnification of pharmaceuticals and endocrine disruptors in a Mediterranean river food web. Science of the Total Environment 540: 250-259.

Boix, D., Caria, M.C., Gascón, S., Mariani, M.A., Sala, J., Ruhi, A., Compte, J. & S. Bagella (2016). Contrasting intra-annual patterns of six biotic groups with different dispersal mode and ability in Mediterranean temporary wetlands. Marine and Freshwater Research 68, 1044-1060.

Sánchez-Montoya, M., von Schiller, D., Ruhi, A., Sting Pechar, G., Proia, L., Miñano, J., Vidal-Abarca, R.R., Suárez, M.L., & K. Tockner (2016). Responses of ground-dwelling arthropods to surface flow drying in channels and adjacent habitats along Mediterranean streams. Ecohydrology 9(7):1376–1387.

Escoriza, D. & A. Ruhi (2016). Functional distance to recipient communities may favour invasiveness: insights from two invasive frogs. Diversity and Distributions 22(5):519-533.

Gascón, S., Arranz, I. Cañedo-Argüelles, M., Nebra, A., Ruhi, A., Rieradevall, M., Caiola, N., Sala, J., Ibàñez, C., Quintana, XD. & D. Boix (2016). Environmental filtering determines metacommunity structure in wetland microcrustaceans. Oecologia 181(1):193-205.

Compte, J., Montenegro, M., Ruhi, A., Gascón, S., Sala, J. & D. Boix (2016). Microhabitat selection and diel patterns of zooplankton in a Mediterranean temporary pond Hydrobiologia 766(1):201-213.

Huerta, B., Rodriguez-Mozaz S., Nannou C., Nakis L., Ruhi A., Acuña V., Sabater, S. & D. Barcelo (2016). Determination of a broad spectrum of pharmaceuticals and endocrine disruptors in biofilm from a WWTP-impacted river. Science of the Total Environment. 540:241-249.

Ruhi, A., Fairchild, G.W., Spieles, D.J., Becerra-Jurado, G. & D. Moreno-Mateos (2016). Invertebrates in Created and Restored Wetlands. In: Batzer, D. & D. Boix. Invertebrates in Freshwater Wetlands: An international perspective on their ecology. Springer.

Ruhi, A., Holmes, E., Rinne, J. & J.L. Sabo (2015). Anomalous droughts, not invasion, decrease persistence of native fishes in a desert river. Global Change Biology 21(4):1482-1496. [media coverage]

Arranz, I., Sala, J., Gascón, S., Ruhi, A., Quintana, X.D., Amoedo, J., Martinoy, M. & D Boix (2015). Contribution to the knowledge of the distribution of Chaoborus species (Diptera: Chaoboridae) in NE Iberian Peninsula, with notes on the spatial and temporal segregation among them. Limnetica 34:57-68.

Huerta, B., Jakimska, A., Llorca, M., Ruhi, A., Margaoutidis, G., Acuña, V., Sabater, S., Rodriguez-Mozaz, S. & D. Barceló (2015). Development of an analytical method for the determination of multi-class pharmaceuticals and endocrine disruptors in freshwater invertebrates. Talanta 132:373-381.

Ruhi, A., Chappuis, E., Escoriza, D., Jover, M., Sala, J., Boix, D., Gascón, S. & E. Gacia (2014). Environmental filtering determines community patterns in temporary wetlands – a multi‑taxon approach. Hydrobiologia 723(1):25-39.

Ruhi, A. & D. Batzer (2014). Assessing congruence and surrogacy among wetland macroinvertebrate taxa towards efficiently measuring biodiversity. Wetlands 34(6):1061-1071.

Escoriza, D. & A. Ruhi (2014). Macroecological patterns of amphibian assemblages in the Western Palearctic: Implications for conservation. Biological Conservation 176:252-261.

Rouissi, M., Boix, D., Muller, S.D., Gascón, S., Ruhi, A., Sala, J., Bouattour, A., Ben Haj Jilani, I., Ghrabi-Gammar, Z., Ben Saad-Limam, S. & A. Daoud-Bouattour (2014). Spatio-temporal variability of faunal and floral assemblages in Mediterranean temporary wetlands. Comptes Rendus Biologies 337(12):695-708.

Ruhi, A., Boix, D., Gascón, S., Sala, J. & D. Batzer (2014). Functional and phylogenetic relatedness in temporary wetland invertebrates: current macroecological patterns and implications for future climatic change scenarios. PloS one 8(11):e81739.

Tornés, A. & A. Ruhi (2013). Flow intermittency decreases nestedness and specialization of diatom communities in Mediterranean rivers. Freshwater Biology 58(12):2555-2566.

Batzer, D. & A. Ruhi (2013). Is there a core set of organisms that structure invertebrate assemblages in freshwater wetlands? Freshwater Biology 58(8):1647-1659.

Ruhi, A., Boix, D., Sala, J., Gascón, S. & X.D. Quintana (2013). Nestedness and successional trajectories of macroinvertebrate assemblages in created wetlands. Oecologia 171(2):545-556.

Ruhi, A., Herrmann, J., Gascón, S., Sala, J., Geijer, J. & D. Boix (2012). Change in biological traits and community structure of macroinvertebrates through primary succession in a man-made Swedish wetland. Freshwater Science 31(1):22-37.

Ruhi, A., San Sebastian, O., Feo, C., Franch, M., Gascón, S., Richter-Boix, À., Boix, D. & G. Llorente (2012). Man-made Mediterranean Temporary Ponds as a tool for amphibian conservation. Annales de Limnologie – International Journal of Limnology 48(1):81-93. [Editor’s choice]

Ruhi, A., Herrmann, J., Gascón, S., Sala, J. & D. Boix (2012). How do early successional patterns in man-made wetlands differ between cold temperate and Mediterranean regions? Limnologica 42(4):328-339.

Boix, D., Ruhi, A., Gascón, S. & J. Sala (2012). First record of Holocentropus stagnalis (Albarda, 1874) (Trichoptera: Polycentropodidae) for the Iberian Peninsula. SEA 51:343‒345.

Boix, D., Ruhi, A., Sala, J., Gascón, S., Compte, J. & X.D. Quintana (2010). Invertebrats aquàtics. In: Fraga, P., I. Estaún & E. Cardona (Eds.). Col.lecció Recerca, 15. Basses temporals mediterrànies. Institut Menorquí d’Estudis.

Ruhi, A., D. Boix, J. Sala, S. Gascón & X. D. Quintana (2009). Spatial and temporal patterns of pioneer macrofauna in recently created ponds: taxonomic and functional approaches. Hydrobiologia 634(1):137-151.