Peer-Reviewed Publications (with lab members in bold)

Perrig, P., S. Lambertucci, P. Alarcón, A. D. Middleton, J. Padró, P. Plaza, G. Blanco. J. A. Sánchez-Zapata, J. Donazar; J. N. Pauli. In review. Fine-scale sexual segregation in foraging sites, yet large dietary overlap, in a dimorphic avian scavenger.

Monk, J. D., J. A. Smith, E. Donadio, P. Perrig, J. N. Pauli, O. Bidder, M. Fileni, P Gregorio, O. J. Schmitz, and A. D. Middleton. In revision. Disease outbreak alters the nature of a trophic cascade.

Barker, K. J., W. Xu, A. Van Scoyoc, M. Serota, J. Moravek, A. Shawler, R. Ryan, and A. D. Middleton. In revision. Can lost migrations be restored? Evaluating the re-establishment of long-distance animal movements.

Bidder, O., J. Morales, G. Rickbeil, J. Merkle; R. Fuda, J. Rogerson, B. Scurlock, W. Edwards, E.  Cole, D. McWhirter, A. Courtemanch, S. Dewey, M. Kauffman, D. MacNulty, J. du Toit, Johan, D. Stahler, A. D. Middleton. In revision. Forage senescence and disease status influence elk pregnancy across the Greater Yellowstone Ecosystem.

Rodriguez Curras, M., E. Donadio, A.D. Middleton, and J.N. Pauli. In revision. Humans affect every major niche axis in a community of carnivores.

Morrison, T., J. Merkle, J. G. C. Hopcraft, E. O. Aikens, J. Beck, R. Boone, A. Courtemanch, S. Dwinnell, S. Fairbanks, B Griffith, A. D. Middleton, K. Monteith, B. Oates, H. Sawyer, K. Smith, J. Stabach, K. Taylor, and M. J. Kauffman. In revision. Cross-species comparison of drivers of site fidelity in ungulates. Journal of Animal Ecology.

Stoellinger, T., H. J. Albers, A.D. Middleton, and J.F. Shogren. In revision. Where the deer and the antelope play: conserving the big game migrations of the West. Duke Environmental Law and Policy Forum.

Xu, W., K. Barker, A. Shawler, A. Van Scoyoc, H. Sawyer, T. Mueller, C. Andreozzi, O. Bidder, H. Karandikar, S. Mumme, J. Smith, E. Templin, and A.D. Middleton. In press. The plasticity of ungulate migrations in a changing world. Ecology.

Middleton, A.D., T. Stoellinger, H. Karandikar, B. Leonard, H. Doremus, and C. Kremen. In press. Harnessing visitors’ enthusiasm for national parks to advance cooperative landscape conservation. Conservation Science and Practice.

Xu, W., D. Nandintsetseg, V. Hermann, H. Sawyer, and A. D. Middleton. In press. Barrier Behavior Analysis (BaBA) reveals extensive effects of fencing on wide-ranging animals. Journal of Applied Ecology.

Smith, J.A., E. Donadio, O.R. Bidder, J.N. Pauli, M.J. Sheriff, and A.D. Middleton. In press. Where and when to hunt? Decomposing predation success of an ambush carnivore. Ecology.

Aikens, E.O., A. Mysterud, J.A. Merkle, F. Cagnacci, I.M. Rivrud, M. Hebblewhite, M.A. Hurley, W. Peters, S. Bergen, J. De Groeve, S.P.H. Dwinnell, B. Gehr, M. Heurich, A.J.M. Hewison, A. Jarnemo, P. Kjellander, M. Kröschel, A. Licoppe, J.D.C. Linnell, E.H. Merrill, A.D. Middleton, N. Morellet, L. Neufeld, A.C. Ortega, K.L. Parker, L. Pedrotti, K.M. Proffitt, S. Said, H. Sawyer, B.M. Scurlock, J. Signer, P. Stent, P. Šustr, T. Szkorupa, K.L. Monteith, and M.J. Kauffman. 2020. Wave-like patterns of plant phenology determine ungulate movement tactics. Current Biology 30: 3444-3449.e4.

Perrig, P.L., S.A. Lambertucci, J. Cruz Bernal, P.A.E. Alarcón, P. Plaza, A.D. Middleton, G. Blanco, J.A. Sánchez, Zapata, J.A. Donazar, and J.N. Pauli. 2020. A regional analysis of habitat selection identifies conservation priority areas for the Andean condor. Biological Conservation 243: 108494.

Middleton, A.D., H. Sawyer, J.A. Merkle, M.J. Kauffman, E.K. Cole, S.R. Dewey, J.A. Gude, D.D. Gustine, D.E. McWhirter, K.M. Proffitt, and P.J. White. 2020. Conserving ungulate migrations across the Greater Yellowstone Ecosystem requires transboundary science, policy, and management. Frontiers in Ecology and the Environment 18: 83-91.

Sawyer, H., C. W. Lebeau, T. L. MacDonald, W. Xu, and A.D. Middleton. 2019. All routes are not created equal: An ungulate’s choice of migration route can influence its survival. Journal of Applied Ecology 56: 1860-69.

Rickbeil, G.J.M., J.A. Merkle, G. Anderson, M.P. Atwood, J.P. Beckmann, E.K Cole, A.B. Courtemanch, S. Dewey, D.D. Gustine, M.J. Kauffman, D.E. McWhirter, T. Mong, K. Profitt, D.R Stahler, P.J. White, and A.D. Middleton. 2019. Plasticity in elk migration timing is a response to changing environmental conditions. Global Change Biology 25: 2368-2381.

Smith, J.A., E. Donadio, J.N. Pauli, M.J. Sheriff, and A.D. Middleton. 2019. Integrating temporal refugia into landscapes of fear: prey exploit predator downtimes to forage in risky places. Oecologia 189:883-890.

Smith, J.A., E. Donadio, J.N. Pauli, M.J. Sheriff, A.D. Middleton. 2019. Habitat complexity mediates the predator-prey space race. Ecology 100: e02724.

Gaynor, K.M., J.S. Brown, A.D Middleton, M.E. Power, and J.S. Brashares. 2019. Landscapes of fear: spatial patterns of risk perception and response. Trends in Ecology and Evolution 34:355-368.

Sawyer, H., J. Merkle, A.D. Middleton, S.P.H.  Dwinnell, and K. Monteith. 2019Migratory plasticity is not ubiquitous among large herbivores. Journal of Animal Ecology 88:450-460.

van Eeden, L.M., A. Eklund, J.R.B. Miller, J. V. Lopez-Bao., G. Chapron, M.R. Cejtin, M. S. Crowther, C.R. Dickman, J. Frank, M. Krofel, D.W. Macdonald, J. McManus, T.R. Meyer, A.D. Middleton, T.M. Newsome, W.J. Ripple, E. G. Richie, O.J. Schmitz, K. J. Stoner, M. Tourani, A. Treves. 2018. Carnivore conservation needs evidence-based livestock protection.  PLoS Biology 16:e2005577.

Middleton, A.D., J.A. Merkle, D.E. McWhirter, J.G. Cook, R.C. Cook, P.J. White, M.J. Kauffman. 2018. Green-wave surfing influences fitness in a large migratory herbivore. Oikos 127: 1060-1068.

Lukacs, P., M. Hebblewhite, B. Johnson, H. Johnson, M. Kauffman, M. Mitchell, K. Proffitt, P. Zager, J. Brodie, K. Hersey, A. Holland, M. Hurley, S. McCorquodale, A.D. Middleton, M. Nordhagen, J. Nowak, D. Walsh, P.J. White. 2018. Factors influencing elk recruitment across three ecotypes in the western USA. Journal of Wildlife Management 82: 698-710.

Cotterill, G.G., P.C. Cross, A.D. Middleton, J.D. Rogerson, B.M Scurlock, J.T. Toit, JT. 2018. Hidden cost of disease in a free‐ranging ungulate: brucellosis reduces mid‐winter pregnancy in elk. Ecology and Evolution 8: 10733-10742.

Perrig, P., E. Donadio, A.D. Middleton, and J.N. Pauli. 2016. Predation subsidizes an obligate scavenger in the high Andes. Journal of Animal Ecology 54: 846-853.

Miller, J.R.B., K.J. Stoner, M.R. Cejtin, T.K. Meyer, A.D. Middleton, and O.J. Schmitz. 2016. Tools for human-carnivore coexistence: A quantitative assessment of techniques for reducing large carnivore depredation of livestock. Wildlife Society Bulletin 40:806-815.

Sawyer, H., A.D. Middleton, K. Monteith, M. Hayes, and M.J. Kauffman. 2016. The extra mile: Ungulate migration distance alters use of seasonal range and exposure to anthropogenic risk. Ecosphere 7:e01534.

Merkle, J.A., K.L. Monteith, E.O. Aikens, M.M. Hayes, K.R. Hersey, A.D. Middleton, B.A. Oates, H. Sawyer, B.M. Scurlock, and M.J. Kauffman. 2016. Large herbivores surf waves of green-up in spring. Proceedings of the Royal Society of London B 283:20160456.

Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, and K. Gerow. 2016. Native prey distribution and migration mediates wolf predation on domestic livestock in the Greater Yellowstone Ecosystem. Canadian Journal of Zoology 94:291-299.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, R.C. Cook, J.G. Cook, M.D. Jimenez, S.E. Albeke, H. Sawyer, and P.J. White. 2013. Linking antipredator behavior to prey demography reveals the limited risk effects of an actively hunting large carnivore. Ecology Letters 16:1023-1030.

Hand, B.K., S. Chen, N. Anderson,  A. Beja-Pereira, P.C. Cross, M. Ebinger, H. Edwards, R.A. Garrott, M.D. Kardos, M. Kauffman, E.L. Landguth, A.D. Middleton, M. Schwartz, B. Scurlock, P.J. White, P. Zager, and G. Luikart. 2014. Limited maternal gene flow among elk populations in the Greater Yellowstone Ecosystem revealed by mitochondrial DNA. Journal of Fish and Wildlife Management 5:124-132.

Middleton, A.D., T.A. Morrison, J.K. Fortin, M.J. Kauffman, C.T. Robbins, K.M. Proffitt, P.J. White, D.E. McWhirter, T.M. Koel, D. Brimeyer, and W.S. Fairbanks. 2013. Grizzly bear predation links the loss of native trout to the demography of migratory elk in Yellowstone. Proceedings of the Royal Society B – Biological Sciences 280:20130870.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Rejoinder: challenge and opportunity in the study of ungulate migration amid environmental change. Ecology 94:1280-86.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Animal migration amid shifting patterns of predation and phenology: Lessons from a Yellowstone elk herd. Ecology 94:1245-56.

Sawyer, H. M.J. Kauffman, A.D. Middleton, T.A. Morrison, R.M. Nielson, and T.B. Wyckoff. 2013. A framework for understanding barrier effects on migratory ungulates. Journal of Applied Ecology 50:68-78.

Christianson, D., R.W. Klaver, A.D. Middleton, and M.J. Kauffman. 2013. Confounded winter and spring conditions on temperate herbivore ranges. Landscape Ecology 28:427-437.

Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, J.J. Barber, and K. Gerow. 2012. Elk migration patterns and human activity influence wolf habitat selection in the Greater Yellowstone Ecosystem. Ecological Applications 22:2293-2307.

Cross, P.C., E.K. Cole, A.P. Dobson, W.H. Edwards, K.L. Hamlin, G. Luikart, A.D. Middleton, B.M. Scurlock, and P.J. White. 2010. Probable causes of increasing elk brucellosis in the Greater Yellowstone Ecosystem. Ecological Applications 20:278-288.

Pauli, J.N., J.P. Whiteman, M. Riley, and A.D. Middleton. 2009. Defining “noninvasive” for sampling of vertebrates. Conservation Biology 24:349-352.

Hohl, A., C. Picard, S.G. Clark, and A.D. Middleton. 2010. Approaches to large-scale conservation: a survey. In Large Scale Conservation in the Common Interest, eds. S.G. Clark, A. Hohl, C. Picard, and D. Newsome. Springer. 191 pp.

Other Publications

Middleton, A.D. 2019. Tracking Wolves, pp. 179-187 in “Matthew Barney, Redoubt” by Yale University Art Gallery, Yale University Press, New Haven, CT. 368 pp.

Middleton, A.D. 2018. Protecting America’s last great animal migrations. The New York Times (op-ed).

Middleton, A.D. 2017. Sustaining migrations in the modern West. In “Yellowstone Migrations by J. Riis. Pp 119-124.

Middleton, A.D. and L. Allison. 2016. Beyond boundaries in the Greater Yellowstone Ecosystem: scientists and ranchers exploring solutions for the future. Western Landowners Alliance. 28pp.

Middleton, A.D. 2016. Seeing Yellowstone: Science at its limits in a conservation crucible. pp. 36-51 in “Invisible Boundaries: Exploring Yellowstone’s Great Animal Migrations” ed. Karen B. McWhorter. Buffalo Bill Center of the West, Cody, WY. 80 pp.

Middleton, A.D. 2016. “Putting the prey back in predator-prey interactions” (Book review). Quarterly Review of Biology 91: 366.

Middleton, A.D. 2015. “Living with large carnivores: do ecology and conservation biology have the answers? (Book review). Ecology 96: 880-81.

Middleton, A.D. 2014. “Changes in habitat quality and predation shape a Yellowstone elk migration” and “Cascading consequences of the lake trout invasion on Yellowstone’s migratory elk?” Yellowstone Science 22:36-37.

Martínez del Rio, C. and A.D. Middleton. 2010. Laws for ecology? (Book review). Ecology 91:1244-1245.